In the Miocene and Pliocene fossil shell beds of the eastern United States, the single most spectacular molluscan species radiation is seen in the ecphora shells (the Tribe Ecphorini). These bizarrely shaped gastropods, with their distinctive ribbed shell sculpture, represent a separate branch of the Subfamily Ocenebridae, Family Muricidae. Characteristically, these muricid gastropods are heavily ornamented with spiral ribs and cords and are considered...