Loss of biodiversity on tropical and subtropical oceanic islands is one of the most pressing conservation issues. These oceanic islands are well known for their unique fauna and ? ora, which evolved over long periods in isolation from external perturbation. However, the maj- ity of these islands in the Paci? c were eventually settled by Polynesians and then by Europeans; by about 200 years ago, only a few island groups remained untouched. The Bonin...