Buds scaly; leaves evergreen and persistent for several years (except in Larix), scattered along the twigs, spirally arranged or tufted, linear, needle-shaped, or scale-like; sterile and fertile flowers separate upon the same plant; stamens (subtended by scales) spirally arranged upon a central axis, each bearing two pollen-sacs surmounted by a broad-toothed connective; fertile flowers composed of spirally arranged bracts or cover-scales, each bract subtending an ovuliferous scale; cover-scale and ovuliferous scale attached at their bases; cover-scale usually remaining small, ovuliferous scale enlarging, especially after fertilization, gradually becoming woody or leathery and bearing two ovules at its base; cones maturing (except in Pinus) the first year; ovuliferous scales in fruit usually known as cone-scales; seeds winged; roots mostly spreading horizontally at a short distance below the surface.
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